Let's repeat the investigation of [czHP] strings, but including the `8' letter:

    cat bio-j-jsa-gut.wds \
      | sed \
          -e 's/^/_/g' \
          -e 's/$/_/g' \
          -e 's/[ql]j/H/' \
          -e 's/[ql]g/P/' \
          -e 's/cs/z/g' \
          -e 's/ij/k/g' \
          -e 's/ix/e/g' \
          -e 's/is/r/g' \
          -e 's/iiu/n/g' \
          -e 's/y/i/g' \
          -e 's/ci/a/g' \
          -e 's/cg/8/g' \
      | enum-contexts -vPAT='[czHP8][czHP8]*' -vCTX=0 \
      | wfreq

       793 0.19 H
       382 0.09 8
       374 0.09 Hc8
       314 0.07 Hcc8
       305 0.07 ccc8
       277 0.06 zcc8
       178 0.04 Hcc
       163 0.04 ccc
       152 0.04 z
       140 0.03 zcc
       102 0.02 Hc
        74 0.02 cc
        56 0.01 cccHc
        49 0.01 zccc8
        49 0.01 Pccc8
        49 0.01 P
        48 0.01 cc8
        46 0.01 zc
        41 0.01 ccccHc
        40 0.01 cccc
        39 0.01 zccHc
        35 0.01 zcccHc
        35 0.01 Hccc8
        34 0.01 zccc
        27 0.01 cccc8
        25 0.01 Hccc
        24 0.01 zccH
        20 0.00 zc8
        18 0.00 cccH
        18 0.00 8zcc8
        16 0.00 Hzcc
        15 0.00 Hzcc8
        14 0.00 cHc
        14 0.00 Pccc
        13 0.00 zcccH
        12 0.00 ccccH
        12 0.00 cHcc
        11 0.00 cccz
        11 0.00 ccH
        11 0.00 8ccc8
         9 0.00 zccHcc
         9 0.00 cccHc8
         8 0.00 cHcc8
         7 0.00 cccHcc8
         7 0.00 Pzcc8
         7 0.00 Hzc8
         6 0.00 zccHcc8
         6 0.00 8cc
         5 0.00 zcccHcc8
         5 0.00 zcH
         5 0.00 ccccz
         5 0.00 cHc8
         5 0.00 c
         5 0.00 Pcc8
         5 0.00 8cc8
         4 0.00 zzcc8
         4 0.00 ccccHcc
         4 0.00 cccHcc
         4 0.00 cH
         4 0.00 Pcc
         4 0.00 Hcccc
         4 0.00 8ccc
         3 0.00 zcccz
         3 0.00 cccP
         3 0.00 ccHc8
         3 0.00 cPcc
         3 0.00 cPc
         3 0.00 P8
         3 0.00 8zcc
         3 0.00 8zc
         2 0.00 zzcc
         2 0.00 zcccPc
         2 0.00 zcccHcc
         2 0.00 zcccHc8
         2 0.00 zccPcc
         2 0.00 zccHc8
         2 0.00 ccz
         2 0.00 ccccHcc8
         2 0.00 cccPcc8
         2 0.00 cccPcc
         2 0.00 cP
         2 0.00 Pzc8
         2 0.00 Pzc
         2 0.00 Pcccc
         2 0.00 Hzc
         2 0.00 Hczcc
         2 0.00 Hczc
         2 0.00 Hccz
         2 0.00 Hcccc8
         2 0.00 H8
         2 0.00 8zccc8
         2 0.00 8zccc
         2 0.00 8cccc
         2 0.00 8c8
         2 0.00 8Hc8
         1 0.00 zzcccHc
         1 0.00 zzccH
         1 0.00 zzcHcc8
         1 0.00 zcz8
         1 0.00 zccz
         1 0.00 zccccHcc
         1 0.00 zcccc
         1 0.00 zcccHcc8cc
         1 0.00 zccPccc8
         1 0.00 zccP
         1 0.00 zccHccc
         1 0.00 zcHcc
         1 0.00 zcHc
         1 0.00 zPcc
         1 0.00 zHcc
         1 0.00 zH
         1 0.00 ccccc
         1 0.00 ccccPcc8
         1 0.00 cccPccc8
         1 0.00 cccHccc8
         1 0.00 ccc8cc
         1 0.00 ccPzccc8
         1 0.00 ccPzccc
         1 0.00 ccPccc8
         1 0.00 ccP
         1 0.00 ccHcc8
         1 0.00 ccHcc
         1 0.00 cc8cc
         1 0.00 cPccc8
         1 0.00 cPccc
         1 0.00 cPcc8
         1 0.00 cPc8
         1 0.00 cHccz
         1 0.00 cHccc8
         1 0.00 Pzcc
         1 0.00 Hczc8
         1 0.00 Hcz8
         1 0.00 Hcz
         1 0.00 Hccz8
         1 0.00 Hc8zcc8
         1 0.00 Hc8cc
         1 0.00 Hc8c8
         1 0.00 Hc8c
         1 0.00 8zcccz
         1 0.00 8zc8
         1 0.00 8cccc8
         1 0.00 8cccHcc8
         1 0.00 8Hzcc
         1 0.00 8Hcc
         1 0.00 88
     ----- ---- ----
      4282 1.00 TOT

  Apparently the `8' (\cg/) does not tend to be surrounded by [czHP] strokes, it is either preceded
  or followed by them.  Thus `8' seems quite unlike `P'.
  
  Let's look at some `P' strings and try to find similar words with the `P' replaced by something else:
  
    cat bio-j-jsa-gut.wds \
      | sed \
          -e 's/^/_/g' \
          -e 's/$/_/g' \
          -e 's/[ql]j/H/' \
          -e 's/[ql]g/P/' \
          -e 's/cs/z/g' \
          -e 's/ij/k/g' \
          -e 's/ix/e/g' \
          -e 's/is/r/g' \
          -e 's/iiu/n/g' \
          -e 's/y/i/g' \
          -e 's/ci/a/g' \
          -e 's/cg/8/g' \
      | egrep '[^czHP]P[^czHP]' \
      | wfreq
      
         8 0.15 _Poe_
         2 0.04 _oPoea_
         2 0.04 _oPar_
         2 0.04 _oPa_
         2 0.04 _Poeccc8a_
         1 0.02 _qoPor_
         1 0.02 _qoPoe_
         1 0.02 _qoPa_
         1 0.02 _qoP_
         1 0.02 _qoP8a_
         1 0.02 _qoHoPa_
         1 0.02 _oePocHca_
         1 0.02 _oPor_
         1 0.02 _oPoe_
         1 0.02 _oPaezcc8a_
         1 0.02 _oPaea_
         1 0.02 _oPae_
         1 0.02 _oPaeHain_
         1 0.02 _ePoe_
         1 0.02 _Poin_
         1 0.02 _Poezcca_
         1 0.02 _Poezca_
         1 0.02 _Poezc8ae_
         1 0.02 _Poezc8a_
         1 0.02 _Poeccc8_
         1 0.02 _Poecca_
         1 0.02 _Poecc8arcn_
         1 0.02 _Poecc8a_
         1 0.02 _Poearar_
         1 0.02 _Poeain_
         1 0.02 _Poeaecc8a_
         1 0.02 _PoeHczcoe_
         1 0.02 _PoeHcca_
         1 0.02 _Poe8zcc8a_
         1 0.02 _Poe8aHa_
         1 0.02 _PoHan_
         1 0.02 _Par_
         1 0.02 _PaHc8a_
         1 0.02 _P8oe_
         1 0.02 _P8aezcor_
         1 0.02 _HoePa_
     ----- ---- ----
        52 1.00 TOT

    set noglob
    foreach f ( \
      '_'.'oe_'  \
      '_o'.'oea_' \
      '_o'.'ar_' \
      '_o'.'a_' \
      '_'.'oeccc8a_' \
    )
      echo " "
      echo "-----------------------------------------------------------------------"
      echo " "
      cat bio-j-jsa-gut.wds \
        | sed \
            -e 's/^/_/g' \
            -e 's/$/_/g' \
            -e 's/[ql]j/H/' \
            -e 's/[ql]g/P/' \
            -e 's/cs/z/g' \
            -e 's/ij/k/g' \
            -e 's/ix/e/g' \
            -e 's/is/r/g' \
            -e 's/iiu/n/g' \
            -e 's/y/i/g' \
            -e 's/ci/a/g' \
            -e 's/cg/8/g' \
        | compare-contexts -rctx 0 -lctx 0 -colw 24 \
            "${f:r}P${f:e}" "${f:r}[^P]${f:e}"  "${f:r}[^P][^P]${f:e}"
    end
    unset noglob
      
    -----------------------------------------------------------------------

         8 1.00 _Poe_             81 0.52 _qoe_             11 0.19 _zcoe_
     ----- ---- ----              25 0.16 _zoe_             11 0.19 _oHoe_
         8 1.00 TOT               17 0.11 _eoe_             11 0.19 _ccoe_
                                  17 0.11 _8oe_              8 0.14 _oeoe_
                                   9 0.06 _roe_              5 0.09 _oroe_
                                   6 0.04 _Hoe_              3 0.05 _Hcoe_
                               ----- ---- ----               2 0.04 _aroe_
                                 155 1.00 TOT                1 0.02 _qooe_
                                                             1 0.02 _oqoe_
                                                             1 0.02 _eHoe_
                                                             1 0.02 _e8oe_
                                                             1 0.02 _aeoe_
                                                             1 0.02 _8roe_
                                                         ----- ---- ----
                                                            57 1.00 TOT

    -----------------------------------------------------------------------

         2 1.00 _oPoea_            2 1.00 _oroea_        ----- ---- ----
     ----- ---- ----           ----- ---- ----               0 1.00 TOT
         2 1.00 TOT                2 1.00 TOT           

    -----------------------------------------------------------------------

         2 1.00 _oPar_            35 0.92 _oHar_             7 1.00 _oeHar_
     ----- ---- ----               1 0.03 _orar_         ----- ---- ----
         2 1.00 TOT                1 0.03 _oear_             7 1.00 TOT
                                   1 0.03 _o8ar_        
                               ----- ---- ----          
                                  38 1.00 TOT           

    -----------------------------------------------------------------------

         2 1.00 _oPa_             25 0.45 _oHa_             21 0.53 _oHca_
     ----- ---- ----              23 0.41 _oea_             10 0.25 _oeHa_
         2 1.00 TOT                6 0.11 _ora_              9 0.23 _oe8a_
                                   2 0.04 _o8a_          ----- ---- ----
                               ----- ---- ----              40 1.00 TOT
                                  56 1.00 TOT           

    -----------------------------------------------------------------------

         2 1.00 _Poeccc8a_         6 0.67 _qoeccc8a_     ----- ---- ----
     ----- ---- ----               2 0.22 _zoeccc8a_         0 1.00 TOT
         2 1.00 TOT                1 0.11 _8oeccc8a_    
                               ----- ---- ----          
                                   9 1.00 TOT           

    -----------------------------------------------------------------------

  It ssems that isolated `P' = {\lg/,\qg/} is closely related to `r'=\is/, 
  `q' = \q/, `z' = \cs/, `8' = \cg/, `e' = \ix/, `H' = {\lj/,\qj/}.
  
    cat bio-j-jsa-gut.wds \
      | sed \
          -e 's/^/_/g' \
          -e 's/$/_/g' \
          -e 's/[ql]j/H/' \
          -e 's/[ql]g/P/' \
          -e 's/cs/z/g' \
          -e 's/ij/k/g' \
          -e 's/ix/e/g' \
          -e 's/is/r/g' \
          -e 's/iiu/n/g' \
          -e 's/y/i/g' \
          -e 's/ci/a/g' \
          -e 's/cg/8/g' \
      | egrep '[^czHP]Pccc[^czHP]' \
      | wfreq
      
        14 0.22 _oPccc8a_
        14 0.22 _Pccc8a_
         8 0.13 _qoPccc8a_
         4 0.06 _oePccc8a_
         4 0.06 _oPccca_
         4 0.06 _Pcccoe_
         4 0.06 _Pccc8ar_
         3 0.05 _Pccca_
         2 0.03 _qoPccca_
         1 0.02 _oqoPccc8a_
         1 0.02 _ePccc8a_
         1 0.02 _aPccca_
         1 0.02 _aPccc8a_
         1 0.02 _Pccc8ae_
         1 0.02 _8oePccc8a_
     ----- ---- ----
        63 1.00 TOT
      
    set noglob
    foreach f ( \
      '_o'.'ccc8a_' \
      '_'.'ccc8a_' \
      '_qo'.'ccc8a_' \
      '_oe'.'ccc8a_' \
      '_o'.'ccca_' \
      '_'.'cccoe_' \
      '_'.'ccc8ar_' \
      '_'.'ccca_' \
      '_qo'.'ccca_' \
    )
      echo " "
      echo "-----------------------------------------------------------------------"
      echo " "
      cat bio-j-jsa-gut.wds \
        | sed \
            -e 's/^/_/g' \
            -e 's/$/_/g' \
            -e 's/[ql]j/H/' \
            -e 's/[ql]g/P/' \
            -e 's/cs/z/g' \
            -e 's/ij/k/g' \
            -e 's/ix/e/g' \
            -e 's/is/r/g' \
            -e 's/iiu/n/g' \
            -e 's/y/i/g' \
            -e 's/ci/a/g' \
            -e 's/cg/8/g' \
        | compare-contexts -rctx 0 -lctx 0 -colw 24 \
            "${f:r}P${f:e}" "${f:r}[^P]${f:e}"  "${f:r}[^P][^P]${f:e}"
    end
    unset noglob

    -----------------------------------------------------------------------

        14 1.00 _oPccc8a_         23 0.79 _oeccc8a_          1 0.50 _orzccc8a_
     ----- ---- ----               5 0.17 _oHccc8a_          1 0.50 _oecccc8a_
        14 1.00 TOT                1 0.03 _o8ccc8a_      ----- ---- ----
                               ----- ---- ----               2 1.00 TOT
                                  29 1.00 TOT           

    -----------------------------------------------------------------------

        14 1.00 _Pccc8a_          52 0.37 _eccc8a_          23 0.49 _oeccc8a_
     ----- ---- ----              36 0.26 _zccc8a_           5 0.11 _oHccc8a_
        14 1.00 TOT               19 0.13 _cccc8a_           4 0.09 _ezccc8a_
                                  14 0.10 _Hccc8a_           2 0.04 _rzccc8a_
                                   9 0.06 _8ccc8a_           2 0.04 _qoccc8a_
                                   5 0.04 _rccc8a_           2 0.04 _ecccc8a_
                                   4 0.03 _accc8a_           2 0.04 _eHccc8a_
                                   1 0.01 _qccc8a_           1 0.02 _o8ccc8a_
                                   1 0.01 _occc8a_           1 0.02 _eoccc8a_
                               ----- ---- ----               1 0.02 _azccc8a_
                                 141 1.00 TOT                1 0.02 _aeccc8a_
                                                             1 0.02 _acccc8a_
                                                             1 0.02 _8zccc8a_
                                                             1 0.02 _8cccc8a_
                                                         ----- ---- ----
                                                            47 1.00 TOT

    -----------------------------------------------------------------------

         8 1.00 _qoPccc8a_        11 0.65 _qoHccc8a_         2 0.40 _qoezccc8a_
     ----- ---- ----               6 0.35 _qoeccc8a_         2 0.40 _qoHcccc8a_
         8 1.00 TOT            ----- ---- ----               1 0.20 _qoecccc8a_
                                  17 1.00 TOT            ----- ---- ----
                                                             5 1.00 TOT

    -----------------------------------------------------------------------

         4 1.00 _oePccc8a_         1 1.00 _oecccc8a_         1 1.00 _oeoHccc8a_
     ----- ---- ----           ----- ---- ----           ----- ---- ----
         4 1.00 TOT                1 1.00 TOT                1 1.00 TOT

    -----------------------------------------------------------------------

         4 1.00 _oPccca_          12 0.63 _oeccca_           2 0.40 _oezccca_
     ----- ---- ----               6 0.32 _oHccca_           2 0.40 _oecccca_
         4 1.00 TOT                1 0.05 _orccca_           1 0.20 _oeHccca_
                               ----- ---- ----           ----- ---- ----
                                  19 1.00 TOT                5 1.00 TOT

    -----------------------------------------------------------------------

         4 1.00 _Pcccoe_           3 0.27 _zcccoe_           1 0.33 _oecccoe_
     ----- ---- ----               3 0.27 _8cccoe_           1 0.33 _cccccoe_
         4 1.00 TOT                2 0.18 _ecccoe_           1 0.33 _8ccccoe_
                                   2 0.18 _Hcccoe_       ----- ---- ----
                                   1 0.09 _acccoe_           3 1.00 TOT
                               ----- ---- ----          
                                  11 1.00 TOT           

    -----------------------------------------------------------------------

         4 1.00 _Pccc8ar_          1 0.25 _eccc8ar_      ----- ---- ----
     ----- ---- ----               1 0.25 _cccc8ar_          0 1.00 TOT
         4 1.00 TOT                1 0.25 _accc8ar_     
                                   1 0.25 _Hccc8ar_     
                               ----- ---- ----          
                                   4 1.00 TOT           

    -----------------------------------------------------------------------

         3 1.00 _Pccca_           31 0.39 _cccca_           12 0.35 _oeccca_
     ----- ---- ----              23 0.29 _zccca_            6 0.18 _oHccca_
         3 1.00 TOT               15 0.19 _eccca_            3 0.09 _ezccca_
                                   5 0.06 _Hccca_            3 0.09 _azccca_
                                   4 0.05 _rccca_            2 0.06 _ecccca_
                                   1 0.01 _accca_            2 0.06 _8zccca_
                                   1 0.01 _8ccca_            1 0.03 _rcccca_
                               ----- ---- ----               1 0.03 _qoccca_
                                  80 1.00 TOT                1 0.03 _orccca_
                                                             1 0.03 _acccca_
                                                             1 0.03 _Hcccca_
                                                             1 0.03 _8cccca_
                                                         ----- ---- ----
                                                            34 1.00 TOT

    -----------------------------------------------------------------------

         2 1.00 _qoPccca_          8 0.50 _qoeccca_          1 0.50 _qoeHccca_
     ----- ---- ----               8 0.50 _qoHccca_          1 0.50 _qoHcccca_
         2 1.00 TOT            ----- ---- ----           ----- ---- ----
                                  16 1.00 TOT                2 1.00 TOT

  It seems that `Pccc' is closely related to `Hccc' `eccc' `zccc' `8ccc' `cccc'. 

    cat bio-j-jsa-gut.wds \
      | sed \
          -e 's/^/_/g' \
          -e 's/$/_/g' \
          -e 's/[ql]j/H/' \
          -e 's/[ql]g/P/' \
          -e 's/cs/z/g' \
          -e 's/ij/k/g' \
          -e 's/ix/e/g' \
          -e 's/is/r/g' \
          -e 's/iiu/n/g' \
          -e 's/y/i/g' \
          -e 's/ci/a/g' \
          -e 's/cg/8/g' \
      | egrep '[^czHP]Pzcc[^czHP]' \
      | wfreq

         3 0.38 _qoPzcc8a_
         2 0.25 _oPzcc8a_
         1 0.12 _oPzcc8ae_
         1 0.12 _oPzcc8_
         1 0.12 _Pzccoe8a_
     ----- ---- ----
         8 1.00 TOT

    set noglob
    foreach f ( \
      '_qo'.'zcc8a_' \
      '_o'.'zcc8a_' \
      '_o'.'zcc8ae_' \
      '_o'.'zcc8_' \
      '_'.'zccoe8a_' \
    )
      echo " "
      echo "-----------------------------------------------------------------------"
      echo " "
      cat bio-j-jsa-gut.wds \
        | sed \
            -e 's/^/_/g' \
            -e 's/$/_/g' \
            -e 's/[ql]j/H/' \
            -e 's/[ql]g/P/' \
            -e 's/cs/z/g' \
            -e 's/ij/k/g' \
            -e 's/ix/e/g' \
            -e 's/is/r/g' \
            -e 's/iiu/n/g' \
            -e 's/y/i/g' \
            -e 's/ci/a/g' \
            -e 's/cg/8/g' \
        | compare-contexts -rctx 0 -lctx 0 -colw 24 \
            "${f:r}P${f:e}" "${f:r}[^P]${f:e}"  "${f:r}[^P][^P]${f:e}"
    end
    unset noglob

    -----------------------------------------------------------------------

         3 1.00 _qoPzcc8a_         7 0.88 _qoHzcc8a_     ----- ---- ----
     ----- ---- ----               1 0.12 _qoezcc8a_         0 1.00 TOT
         3 1.00 TOT            ----- ---- ----          
                                   8 1.00 TOT           

    -----------------------------------------------------------------------

         2 1.00 _oPzcc8a_         14 0.88 _oezcc8a_          1 1.00 _oeHzcc8a_
     ----- ---- ----               2 0.12 _oHzcc8a_      ----- ---- ----
         2 1.00 TOT            ----- ---- ----               1 1.00 TOT
                                  16 1.00 TOT           

    -----------------------------------------------------------------------

         1 1.00 _oPzcc8ae_     ----- ---- ----           ----- ---- ----
     ----- ---- ----               0 1.00 TOT                0 1.00 TOT
         1 1.00 TOT                                     

    -----------------------------------------------------------------------

         1 1.00 _oPzcc8_           2 1.00 _oezcc8_       ----- ---- ----
     ----- ---- ----           ----- ---- ----               0 1.00 TOT
         1 1.00 TOT                2 1.00 TOT           

    -----------------------------------------------------------------------

         1 1.00 _Pzccoe8a_     ----- ---- ----           ----- ---- ----
     ----- ---- ----               0 1.00 TOT                0 1.00 TOT
         1 1.00 TOT                                     

  Again the `P' seems to be similar to `H' and `e'.
  
  And now for something completely different.  Let's look at how the words are distributed among the 
  paragraphs:
  
    cat bio-j-jsa.wds \
        | sed \
            -e 's/[ql]j/H/g' \
            -e 's/[ql]g/P/g' \
            -e 's/ij/k/g' \
            -e 's/ix/e/g' \
            -e 's/is/r/g' \
            -e 's/iiu/n/g' \
            -e 's/cy/a/g' \
            -e 's/ci/a/g' \
            -e 's/in/m/g' \
            -e 's/ir/w/g' \
            -e 's/cs/z/g' \
            -e 's/cg/8/g' \
        | enum-words-in-blocks -vWPB=100 \
        | egrep -v '[^a-zA-Z0-9_ ]' \
        | sort +1 -2 +0 -1n \
        | make-word-location-map -vNBLOCKS=71 \
        > .foo

  The result has been posted as http://www.dcc.unicamp.br/~stolfi/voynich/word-distr-map.html
  
  Recomputing with fewer large blocks:
  
    cat bio-j-jsa.wds \
      | sed \
          -e 's/[ql]j/H/g' \
          -e 's/[ql]g/P/g' \
          -e 's/ij/k/g' \
          -e 's/ix/e/g' \
          -e 's/is/r/g' \
          -e 's/iiu/n/g' \
          -e 's/cy/a/g' \
          -e 's/ci/a/g' \
          -e 's/in/m/g' \
          -e 's/ir/w/g' \
          -e 's/cs/z/g' \
          -e 's/cg/8/g' \
      | enum-words-in-blocks -vWPB=1010 \
      | egrep -v '[^a-zA-Z0-9_ ]' \
      | sort +1 -2 +0 -1n \
      | make-word-location-map -vCTWD=3 -vNBLOCKS=7 \
      > .foo
        
    cat .foo \
      | gawk '/./ { printf"%5d %-16s ", $1, $2; for (i=3; i<=NF; i++) printf " %2d", int(($(i)*99/$1)+0.5); printf "\n" }' \
      > .bar

  Results posted in my Voynich page.